SUP3RN0VA
DR4G0N K1NG
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Yes. They are the physical manifestations of the moon and ocean. Hence why when Tui was killed, the moon disappeared. But their physical forms are that of koi fish. Hence, the 'animal' that represents waterbending is the koi fish.
Although if we really want to get technical, it was the lion turtles who gave humans the power to bend in the first place. But four lion turtle based Starters would get boring.
I am talking about the original benders. Neither Koi Fish were the original bender. The Moon is the original water bender and that is what the Waterbenders learnt from.
The Lion turtles used Energybending to bestow upon humans the ability to control the elements, and it was the Moon, Badger Mole, Dragon and Flying Bison that the humans learnt from to control their powers and turn it into the art form known as 'bending.'